diff --git a/spring-core/src/test/kotlin/org/springframework/core/CoroutinesUtilsTests.kt b/spring-core/src/test/kotlin/org/springframework/core/CoroutinesUtilsTests.kt
index 6e0a13342f6..4cc2fdeefdb 100644
--- a/spring-core/src/test/kotlin/org/springframework/core/CoroutinesUtilsTests.kt
+++ b/spring-core/src/test/kotlin/org/springframework/core/CoroutinesUtilsTests.kt
@@ -230,7 +230,7 @@ class CoroutinesUtilsTests {
 	}
 
 	@Test
-	suspend fun invokeSuspendingFunctionWithNullableValueClassParameterWithNonnullObject() {
+	suspend fun invokeSuspendingFunctionWithNullableValueClassParameterAndNonNullParameter() {
 		val method = CoroutinesUtilsTests::class.java.declaredMethods.first { it.name.startsWith("suspendingFunctionWithNullableValueClass") }
 		val mono = CoroutinesUtils.invokeSuspendingFunction(method, this, ValueClass("foo"), null) as Mono
 		Assertions.assertThat(mono.awaitSingleOrNull()).isEqualTo("foo")
diff --git a/spring-web/src/main/java/org/springframework/web/method/support/InvocableHandlerMethod.java b/spring-web/src/main/java/org/springframework/web/method/support/InvocableHandlerMethod.java
index 07a4b78790d..6dca729698e 100644
--- a/spring-web/src/main/java/org/springframework/web/method/support/InvocableHandlerMethod.java
+++ b/spring-web/src/main/java/org/springframework/web/method/support/InvocableHandlerMethod.java
@@ -316,7 +316,7 @@ public class InvocableHandlerMethod extends HandlerMethod {
 						Object arg = args[index];
 						if (!(parameter.isOptional() && arg == null)) {
 							KType type = parameter.getType();
-							if (!(type.isMarkedNullable() && arg == null) &&
+							if (!type.isMarkedNullable() &&
 									type.getClassifier() instanceof KClass<?> kClass &&
 									KotlinDetector.isInlineClass(JvmClassMappingKt.getJavaClass(kClass))) {
 								arg = box(kClass, arg);
@@ -337,7 +337,7 @@ public class InvocableHandlerMethod extends HandlerMethod {
 		private static Object box(KClass<?> kClass, @Nullable Object arg) {
 			KFunction<?> constructor = Objects.requireNonNull(KClasses.getPrimaryConstructor(kClass));
 			KType type = constructor.getParameters().get(0).getType();
-			if (!(type.isMarkedNullable() && arg == null) &&
+			if (!type.isMarkedNullable() &&
 					type.getClassifier() instanceof KClass<?> parameterClass &&
 					KotlinDetector.isInlineClass(JvmClassMappingKt.getJavaClass(parameterClass))) {
 				arg = box(parameterClass, arg);
diff --git a/spring-web/src/test/kotlin/org/springframework/web/method/support/InvocableHandlerMethodKotlinTests.kt b/spring-web/src/test/kotlin/org/springframework/web/method/support/InvocableHandlerMethodKotlinTests.kt
index 4b15b492193..b7da2b669f2 100644
--- a/spring-web/src/test/kotlin/org/springframework/web/method/support/InvocableHandlerMethodKotlinTests.kt
+++ b/spring-web/src/test/kotlin/org/springframework/web/method/support/InvocableHandlerMethodKotlinTests.kt
@@ -148,6 +148,13 @@ class InvocableHandlerMethodKotlinTests {
 		Assertions.assertThatIllegalArgumentException().isThrownBy { invocable.invokeForRequest(request, null) }
 	}
 
+	@Test
+	fun valueClassWithNullableAndNonNullParameter() {
+		composite.addResolver(StubArgumentResolver(LongValueClass::class.java, LongValueClass(1)))
+		val value = getInvocable(ValueClassHandler::valueClassWithNullable.javaMethod!!).invokeForRequest(request, null)
+		Assertions.assertThat(value).isEqualTo(1L)
+	}
+
 	@Test
 	fun valueClassWithNullable() {
 		composite.addResolver(StubArgumentResolver(LongValueClass::class.java, null))
diff --git a/spring-webflux/src/main/java/org/springframework/web/reactive/result/method/InvocableHandlerMethod.java b/spring-webflux/src/main/java/org/springframework/web/reactive/result/method/InvocableHandlerMethod.java
index 920e29b5ef8..c06ea62a2aa 100644
--- a/spring-webflux/src/main/java/org/springframework/web/reactive/result/method/InvocableHandlerMethod.java
+++ b/spring-webflux/src/main/java/org/springframework/web/reactive/result/method/InvocableHandlerMethod.java
@@ -329,7 +329,7 @@ public class InvocableHandlerMethod extends HandlerMethod {
 		private static final String COROUTINE_CONTEXT_ATTRIBUTE = "org.springframework.web.server.CoWebFilter.context";
 
 		@SuppressWarnings("DataFlowIssue")
-		public static @Nullable Object invokeFunction(Method method, Object target, Object[] args, boolean isSuspendingFunction,
+		public static @Nullable Object invokeFunction(Method method, Object target, @Nullable Object[] args, boolean isSuspendingFunction,
 				ServerWebExchange exchange) throws InvocationTargetException, IllegalAccessException, NoSuchMethodException {
 
 			if (isSuspendingFunction) {
@@ -356,7 +356,7 @@ public class InvocableHandlerMethod extends HandlerMethod {
 							Object arg = args[index];
 							if (!(parameter.isOptional() && arg == null)) {
 								KType type = parameter.getType();
-								if (!(type.isMarkedNullable() && arg == null) &&
+								if (!type.isMarkedNullable() &&
 										type.getClassifier() instanceof KClass<?> kClass &&
 										KotlinDetector.isInlineClass(JvmClassMappingKt.getJavaClass(kClass))) {
 									arg = box(kClass, arg);
@@ -378,7 +378,7 @@ public class InvocableHandlerMethod extends HandlerMethod {
 		private static Object box(KClass<?> kClass, @Nullable Object arg) {
 			KFunction<?> constructor = Objects.requireNonNull(KClasses.getPrimaryConstructor(kClass));
 			KType type = constructor.getParameters().get(0).getType();
-			if (!(type.isMarkedNullable() && arg == null) &&
+			if (!type.isMarkedNullable() &&
 					type.getClassifier() instanceof KClass<?> parameterClass &&
 					KotlinDetector.isInlineClass(JvmClassMappingKt.getJavaClass(parameterClass))) {
 				arg = box(parameterClass, arg);
diff --git a/spring-webflux/src/test/kotlin/org/springframework/web/reactive/result/InvocableHandlerMethodKotlinTests.kt b/spring-webflux/src/test/kotlin/org/springframework/web/reactive/result/InvocableHandlerMethodKotlinTests.kt
index 95a5df9cbbc..9e41a222399 100644
--- a/spring-webflux/src/test/kotlin/org/springframework/web/reactive/result/InvocableHandlerMethodKotlinTests.kt
+++ b/spring-webflux/src/test/kotlin/org/springframework/web/reactive/result/InvocableHandlerMethodKotlinTests.kt
@@ -250,6 +250,14 @@ class InvocableHandlerMethodKotlinTests {
 		assertExceptionThrown(result, IllegalArgumentException::class)
 	}
 
+	@Test
+	fun valueClassWithNullableAndNonNullParameter() {
+		this.resolvers.add(stubResolver(LongValueClass(1), LongValueClass::class.java))
+		val method = ValueClassController::valueClassWithNullable.javaMethod!!
+		val result = invoke(ValueClassController(), method)
+		assertHandlerResultValue(result, "1")
+	}
+
 	@Test
 	fun valueClassWithNullable() {
 		this.resolvers.add(stubResolver(null, LongValueClass::class.java))